Tim Weaver

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​I study human evolution, with a focus on the origins, evolution, and disappearance of Neandertals, and the related topic of the origins of humans who were anatomically and behaviorally modern. While my emphasis is on the later phases of human evolution, I have also worked on earlier periods, mostly in the context of the evolution of human bipedal walking and running and childbirth.

Most of my publications address specific questions about why Neandertal, early modern human, or present-day human skeletons look the way they do. In my work, I strive to integrate approaches and datasets from population and molecular genetics with traditional studies of the fossil and archaeological records.

Journal articles

  • Weaver, Timothy D. 2018. Neutral theory and the evolution of human physical form: an introduction to models and applications. Journal of Anthropological Sciences 96:7-26.
  • Henn, Brenna M., Teresa E. Steele, and Timothy D. Weaver. 2018. Clarifying distinct models of modern human origins in Africa. Current Opinion in Genetics and Development 53:148–156.
  • Weaver, Timothy D. and Philipp Gunz. 2018. Using geometric morphometric visualizations of directional selection gradients to investigate morphological differentiation. Evolution 72:838-850.
  • Katz, David C., Mark N. Grote, and Timothy D. Weaver. 2017. Changes in human skull morphology across the agricultural transition are consistent with softer diets in pre-industrial farming groups. Proceedings of the National Academy of Sciences of the U.S.A. 114:9050-9055.
  • Weaver, Timothy D. 2016. Estimators for QST and coalescence times. Ecology and Evolution 6:7783-7786.
  • Katz, David C., Mark N. Grote, and Timothy D. Weaver. 2016. A mixed model for the relationship between climate and human cranial form. American Journal of Physical Anthropology 160:593-603.
  • Weaver, Timothy D., Hélène Coqueugniot, Luibov V. Golovanova, Vladimir B. Doronichev, Bruno Maureille, and Jean-Jacques Hublin. 2016. Neonatal postcrania from Mezmaiskaya, Russia, and Le Moustier, France, and the development of Neandertal body form. Proceedings of the National Academy of Sciences of the U.S.A. 113:6472-6477.
  • Gilmore, Cassandra C. and Timothy D. Weaver. 2016. Comparative perspective on antemortem tooth loss in Neandertals. Journal of Human Evolution 92:80-90.
  • Weaver, Timothy D. and Chris B. Stringer. 2015. Unconstrained cranial evolution in Neandertals and modern humans compared to common chimpanzees. Proceedings of the Royal Society B: Biological Sciences 282:20151519.
  • Guo, Jing, Jingze Tan, Yajun Yang, Hang Zhou, Sile Hu, Agu Hashan, Nurmamat Bahaxar, Shuhua Xu, Timothy D. Weaver, Li Jun, Mark Stoneking, and Kun Tang. 2014. Variation and signatures of selection on the human face. Journal of Human Evolution 75:143-152.
  • Weaver, Timothy D. 2014. Brief Communication: quantitative- and molecular-genetic differentiation in humans and chimpanzees: implications for the evolutionary processes underlying cranial diversification. American Journal of Physical Anthropology 154:615-620.
  • Weaver, Timothy D. 2014. Tracing the paths of modern humans from Africa. Proceedings of the National Academy of Sciences of the U.S.A. 111:7170-7171.
  • Weaver, Timothy D. 2012. Did a discrete event 200,000-100,000 years ago produce modern humans? Journal of Human Evolution 63:121-126.
  • Steele, Teresa E. and Timothy D. Weaver. 2012. Refining the Quadratic Crown Height Method of age estimation: do elk teeth wear quadratically with age? Journal of Archaeological Science 39:2329-2334.
  • Weaver, Timothy D., Ryan H. Boyko, and Teresa E. Steele. 2011. Cross-platform program for likelihood-based statistical comparisons of mortality profiles on a triangular graph. Journal of Archaeological Science 38:2420-2423.
  • Roseman, Charles C., Timothy D. Weaver, and Chris B. Stringer. 2011. Do modern humans and Neandertals have different patterns of cranial integration? Journal of Human Evolution 60:684-693.
  • Weaver, Timothy D., Teresa E. Steele, and Richard G. Klein. 2011. The abundance of eland, buffalo, and wild pigs in Middle and Later Stone Age sites. Journal of Human Evolution 60:309-314.
  • Coqueugniot, Hélène, Timothy D. Weaver, and Francis Houët. 2010. Brief communication: a probabilistic approach to age estimation from infracranial sequences of maturation. American Journal of Physical Anthropology 142:655-664.
  • von Cramon-Taubadel, Noreen and Timothy D. Weaver. 2009. Insights from a quantitative genetic approach to human morphological evolution. Evolutionary Anthropology 18:237-240.
  • Weaver, Timothy D. 2009. The meaning of Neandertal skeletal morphology. Proceedings of the National Academy of Sciences of the U.S.A. 106:16028-16033 (reprinted in: Companion to human evolution. 2014. McBrearty, Sally, Ed. Cognella, 325-338).
  • Bailey, Shara E., Timothy D. Weaver, and Jean-Jacques Hublin. 2009. Who made the Aurignacian and other early Upper Paleolithic industries? Journal of Human Evolution 57:11-26.
  • Weaver, Timothy D. and Jean-Jacques Hublin. 2009. Neandertal birth canal shape and the evolution of human childbirth. Proceedings of the National Academy of Sciences of the U.S.A. 106:8151-8156.
  • Bailey, Shara, Michelle Glantz, Timothy D. Weaver, and Bence Viola. 2008. The affinity of the dental remains from Obi-Rakhmat Grotto, Uzbekistan. Journal of Human Evolution 55:238-248.
  • Weaver, Timothy D., Charles C. Roseman, and Chris B. Stringer. 2008. Close correspondence between quantitative- and molecular-genetic divergence times for Neandertals and modern humans. Proceedings of the National Academy of Sciences of the U.S.A. 105:4645-4649.
  • Weaver, Timothy D. and Charles C. Roseman. 2008. New developments in the genetic evidence for modern human origins. Evolutionary Anthropology 17:69-80.
  • Roseman, Charles C. and Timothy D. Weaver. 2007. Molecules versus morphology? Not for the human cranium. BioEssays 29:185-188.
  • Coqueugniot, Hélène and Timothy D. Weaver. 2007. Brief communication: infracranial maturation in the skeletal collection from Coimbra, Portugal: new aging standards for epiphyseal union. American Journal of Physical Anthropology 134:424-437.
  • Tilkens, Michael J., Cara Wall-Scheffler, Timothy D. Weaver, and Karen Steudel-Numbers. 2007. The effects of body proportions on thermoregulation: an experimental assessment of Allen’s rule. Journal of Human Evolution 53:286-291.
  • Steudel-Numbers, Karen L., Timothy D. Weaver, and Cara M. Wall-Scheffler. 2007. The evolution of human running: effects of changes in lower-limb length on locomotor economy. Journal of Human Evolution 53:191-196.
  • Weaver, Timothy D., Charles C. Roseman, and Chris B. Stringer. 2007. Were Neandertal and modern human cranial differences produced by natural selection or genetic drift? Journal of Human Evolution 53:135-145.
  • Harvati, Katerina and Timothy D. Weaver. 2006. Human cranial anatomy and the differential preservation of population history and climate signatures. The Anatomical Record Part A 288A:1225-1233.
  • Steudel-Numbers, Karen and Timothy D. Weaver. 2006. Froude number corrections in Anthropological studies. American Journal of Physical Anthropology 131:27-32.
  • Weaver, Timothy D. and Karen Steudel-Numbers. 2005. Does climate or mobility explain the differences in body proportions between Neandertals and their Upper Paleolithic successors? Evolutionary Anthropology 14:218-223.
  • Weaver, Timothy D. and Charles C. Roseman. 2005. Ancient DNA, late Neandertal survival, and modern-human-Neandertal genetic admixture. Current Anthropology 46:677-683.
  • Roseman, Charles C. and Timothy D. Weaver. 2004. Multivariate apportionment of global human craniometric diversity. American Journal of Physical Anthropology 125:257-263.
  • Weaver, Timothy D. 2003. The shape of the Neandertal femur is primarily the consequence of a hyperpolar body form. Proceedings of the National Academy of Sciences of the U.S.A. 100:6926-6929.
  • Steele, Teresa E. and Timothy D. Weaver. 2002. The modified triangular graph: A refined method for comparing mortality profiles in archaeological samples. Journal of Archaeological Science 29:317-322.
  • Weaver, Timothy D., Suhail A. Islam, and David L. Weaver. 1994. Mac Molecular: A program for visualization of molecular structures on the Macintosh. Journal of Molecular Graphics 12:231-234.

Book chapters

  • Bailey, Shara E., Timothy D. Weaver, and Jean-Jacques Hublin. 2017. The dentition of the earliest modern humans: how 'modern' are they? In: Human paleontology and prehistory. Contributions in honor of Yoel Rak. Marom, Assaf and Erella Hovers, Eds. Springer: New York, 215-232.
  • Weaver, Timothy D. 2011. Rates of cranial evolution in Neandertals and modern humans. In: Computational paleontology. Elewa, Ashraf, Ed. Springer: New York, 165-178.
  • Harvati, Katerina and Timothy D. Weaver. 2006. Reliability of cranial morphology in reconstructing Neanderthal phylogeny. In: Neanderthals revisited: new approaches and perspectives. Harvati, Katerina and Terry Harrison, Eds. Springer: New York, 239-254.
  • Weaver, Timothy D. and Richard G. Klein. 2005. The evolution of human walking. In: Human Walking. 3rd Edition. Rose, Jessica and James G. Gamble, Eds. Lippincott, Williams, and Wilkins: Philadelphia, 23-32.

Comments, reviews, theses, and multimedia

  • Weaver, Timothy D. In Press. Book review of “50 great myths of human evolution: understanding misconceptions about our origins”, John H. Relethford. Journal of Anthropological Research.
  • Weaver, Timothy D. 2015. Genetics and human skeletal variation. In: Human population genetics II, the biomedical and life sciences collection, Henry Stewart Talks Ltd, London. Rosenberg, Noah A. and Nielsen, Rasmus, Eds. (Online at: https://hstalks.com/t/2999/genetics-and-human-skeletal-variation/?biosci)
  • Steele, Teresa E. and Timothy D. Weaver. 2014. Comment on Cieri et al. Current Anthropology 55:434-435.
  • Weaver, Timothy D. 2003. Book review of “The speciation of modern Homo sapiens,” Tim J. Crow, Ed. The Quarterly Review of Biology 78:513.
  • Weaver, Timothy D. 2002. A multi-causal functional analysis of hominid hip morphology. Ph.D. dissertation. Stanford University, Stanford, U.S.A. (Online at: http://www.paleoanthro.org/dissertations/download)
  • Weaver, Timothy D. 1998. Examining Plio/Pleistocene hominid subsistence activities from a community ecology perspective. Masters thesis. Stanford University, Stanford, U.S.A.
  • Weaver, Timothy D. 1995. Interactive computer visualization of fossil specimens. Honors thesis. Dartmouth College, Hanover, U.S.A

ResearchGate:
​https://www.researchgate.net/profile/Timothy_Weaver

Academia.edu:
http://ucdavis.academia.edu/TimothyWeaver 

Documents